W. Steven Ward, PhD


Bachelor of Science in Chemistry, 1980
American University of Beirut
Beirut, Lebanon

Master of Science in Biochemistry, 1981
American University of Beirut
Beirut, Lebanon

Doctor of Philosophy in Biochemistry, 1985
Vanderbilt University
Nashville, TN

Postdoctoral Fellowship, 1989
John Hopkins University
Baltimore, MD

FOCUS Research

Lakshmi Devi and Devraj Sharma Endowed Chair, 2015
John A. Burns School of Medicine, University of Hawaii
Department of OB/GYN

Edwin A. Beer Award, 1992
New York Academy of Sciences

Fellow, 1985-1989
John Hopkins Oncology Center


Ward, W. S. (2017). Eight tests for sperm DNA fragmentation and their roles in the clinic. Transl Androl Urol, 6(Suppl 4), S468-S470. doi:10.21037/tau.2017.03.78

Nguyen, H., James, N. G., Nguyen, L., Nguyen, T. P., Vuong, C., Ortega, M. A., . . . Ward, W. S. (2017). Higher Order Oligomerization of the Licensing ORC4 Protein Is Required for Polar Body Extrusion in Murine Meiosis. J Cell Biochem, 118(9), 2941-2949. doi:10.1002/jcb.25949

Ortega, M. A., Ko, M., Marh, J., Finberg, A., Oshiro, M., & Ward, W. S. (2016). Presence of the Paternal Pronucleus Assists Embryo in Overcoming Cycloheximide Induced Abnormalities in Zygotic Mitosis. J Cell Biochem, 117(8), 1806-1812. doi:10.1002/jcb.25480

For a more complete list of publications, click here.

Ward Steve
W. Steven Ward PhD
Chief of Research Division

Institute for Biogenesis Research

Professor of Anatomy, Biochemistry, and Physiology
University of Hawai’i